Zwackhiomacromyces hyalosporus
Anamorph: not known.
Teleomorph: ascomata perithecia, not stromatic, erumpent from host areoles, the host tissues slightly darker and greyer but otherwise apparently unaffected, with about half of the perithecia exposed, black, smooth, subglobose, 180-240 µm diam., with a minute but distinctly papillate ostiole. Peridium dark brown, composed of several layers of subglobose to polyhedral cells, with the innermost layer of flattened, almost hyaline to pale brown cells. Interascal tissue of persistent narrow pseudoparaphyses 1.5-2 µm diam., densely branched and anastomosed, sometimes weakly moniliform, not staining blue in iodine. Asci 130-160 x 26-30 µm, cylindric-clavate, short-stalked, thick-walled and fissitunicate, the apex with a narrow to broad ocular chamber, 8-spored (sometimes with only two or four spores developing fide Alstrup & Hawksworth 1990). Ascospores arranged biseriately, (36-) 37-42.5 x 11-13 µm, cylindric-ellipsoidal to broadly fusiform, sometimes slightly asymmetrical, the apices usually obtuse, mostly 3-septate but with a proportion with one or two septa, slightly to strongly constricted at the septa, hyaline (overmature spores may be pale brown to olivaceous), smooth to very faintly verrucose, thin-walled, lacking a well-defined perispore or appendages.
The distinction between Zwackhiomyces and Zwackhiomacromyces is unclear apart from in ascospore dimensions. Molecular data are lacking.
On thalli of Placopsis lambii, the affected host thallus somewhat darker and greyer than the surrounding tissues.
In GBI, known from a single former mine site in VC46 Cardigan.