S. D.  Leavitt, Esslinger, T. L., Divakar, P. K., and Lumbsch, H. T., ?Miocene and Pliocene dominated diversification of the lichen-forming fungal genus Melanohalea (Parmeliaceae, Ascomycota) and Pleistocene population expansions?, BMC Evolutionary Biology, vol. 12, no. 176, p. 18 pp., 2012.