Tubakia dryina  

Anamorph: conidiomata in necrotic areas on living leaves, pycnothyrial, umbilicate with a central column and a shield-like upper part. Scutellum (shield) 60-110 (-125) µm diam., shallowly convex, composed of a single layer of radiating irregular septate thick-walled brown hyphae, with closely appressed branches in the outer part and the apices ± acute. Column composed of several appressed thick-walled pale brown vertically oriented hyphal cells, with a dense short palisade of conidiogenous cells formed from the exposed surface. Conidiogenous cells rather irregular, conical to broadly cylindrical, hyaline, thin-walled, proliferating percurrently with inconspicuous periclinal thickening and no collarette, sometimes forming a small number of annellations. Conidia 10-15 x 7-10.5 µm, obovoid to ellipsoidal, initially hyaline but becoming brown, thick-walled, aseptate, smooth to finely verrucose. Microconidia sometimes produced from the same pycnothyrium as macroconidia or from smaller pycnothyria, 4-9 x 1.5-2.5 µm, fusiform, hyaline. Conidiomata on twigs 200-1000 µm diam., irregular in form and sometimes coalescing, with a black crustose outer wall dehiscing by irregular fissures. Conidia tend to be somewhat narrower than those from leaves, mostly 6–7.5 µm in width.
Teleomorph: stromata in overwintered leaves, 220-280 (-300) µm diam., 200-250 µm tall, mostly scattered, occasionally in small clusters and then sometimes coalesced, amphigenous, containing a single ascoma with a very inconspicuous, usually epiphyllous periphysate ostiole and no distinct neck; shining black. Stroma composed of dark brown, thick-walled and sometimes occluded, angular to rounded cells, the peridium poorly defined and composed of a few layers of thin-walled pale flattened tissue. Interascal tissue absent. Asci 57-63 x 10-12 µm, cylindric-clavate to cylindric-ellipsoidal, short-stalked, thin-walled, the apex rounded to truncate with a very conspicuous obconic-truncate refractive ring 4-5 µm diam. and ca 3 µm tall, eventually deliquescing at the base, 8-spored. Ascospores 11.5-13 x 4-5 µm, ovoid to ovoid-cylindrical, often slightly inaequilateral, aseptate, thin-walled, smooth, hyaline, the contents granular, without a gelatinous sheath or appendages.