Spermogonia 90-130 µm diam., hyphophyllous, subepidermal, mixed with the aecia, yellow, with projecting paraphyses. Aecia to 30 mm in length, chiefly on the swollen nerves and petioles, dark cinnamon brown. Aeciospores similar to the urediniospores. Uredinia hypophyllous, rarely epiphyllous, scattered, minute, punctiform, brown. Urediniospores 25-50 x 18-28 µm, subglobose to obovoid, the wall ca 2 µm thick laterally and 5-6 µm at the apex, brown, regularly and distantly echinulate, distance between spines 3-3.5 µm, with 3-4 ± equatorial pores with swollen caps. Telia scattered, hypophyllous or on the stems, minute, often confluent, long covered by the host epidermis, blackish brown. Teliospores variable in shape with a broad length/breadth amplitude, 30-45 x 18-24 µm, cylindrical to obovoid, sometimes distinctly and irregularly angular, thick-walled but not markedly more so at the apex, hardly constricted at the ± median septum, the upper cell with an ± apical pore and the lower with a pore that is usually near the base, pedicels short, rather broad, hyaline, deciduous.
Description mostly adapted from Wilson & Henderson (1966).