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Helvella monachella
Nomenclature
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Family: HelvellaceaeGenus: Helvella
SUMMARY
Anamorph: not known.
Teleomorph: stromata absent. Ascomata discomycetous, stipitate. Cap (10-) 25-35 mm diam. and 17-40 mm high, saddle-shaped or mitre-like with two or three distinct irregular lobes. Hymenium brown-black to black (black when dry), sometimes with purple or chocolate brown tinges. Outer surface white to pale grey, glabrous. Ectal excipulum (outer surface layer) 85-150 µm thick, composed of chains of angular hyaline cells, the outermost cells cylindrical to clavate, 20-60 x 6-20 µm in size. Internal tissue (below the hymenium) 200-300 µm thick, composed of interwoven hyphae 3-6 µm diam., interspersed with thicker-walled hyphae 6-10 µm diam. Stipe 20-55 × 7-10 mm, generally squat, often expanded at the base (to 15-20 mm), round in section, sometimes slightly furrowed below, hollow, white, sometimes stained pale grey or brown with age, glabrous, which may be partly buried. Interascal tissue of unbranched paraphyses with swollen apices to 5-9 µm diam., containing a brown pigment. Asci 310-360 × 14-17 µm, cylindrical, fairly thick-walled, the apex rounded, operculate, 8-spored. Ascospores 20-24 × 13-15 (-16) µm, mean length/breadth ratio 1.5, ellipsoidal to broadly ellipsoidal, fairly thin-walled, hyaline, smooth, containing a large central lipid droplet surrounded by smaller ones, without a gelatinous sheath or appendages.
Description adapted from Dissing (1966) and van Vooren (2010). Formerly known as Helvella leucopus, but H. monachella is an older name according to Skrede et al. (2017).