Thallus squamulose. Squamules to 4 mm diam., widely scattered or contiguous, rounded or irregular. Upper surface flat or irregularly convex, pale grey to dark brown, often tinged green, pruinose or not, dull, usually with faint, paler spots and short lines along more convex areas and ridges, without regular fissures and pseudocyphellae, margin concolorous with upper side, not pruinose. Upper cortex 60-90 μm thick, lacking a well-defined epinecral layer, without calcium oxalate crystals. Lower cortex poorly developed or absent. Medulla lacking crystals. Photobiont zone continuous, containing chlorococcoid algae. Anamorph: conidiomata pycnidia, immersed in the squamules. Conidia filiform. Teleomorph: ascomata apothecia, to 1.5 mm diam., ± persistently plane and marginate, not or faintly white-pruinose, often irregular in form. True exciple raised at first but often becoming excluded, confluent with the hypothecium, uniformly dark reddish brown, K–, N–. Epithecium olivaceous green to bright green, lacking crystals, K–, N+ violet. Hymenium 70-80 μm tall, hyaline to pale brown. Hypothecium dark reddish brown. Interascal tissue of paraphyses, straight, simple or sparingly branched and anastomosed, not conglutinated, thin-walled, the apical cell distinctly swollen and covered by a ± well-developed, gelatinous pigment cap. Asci clavate, surrounded by a gelatinous, K/I+ blue sheath, with a well-developed, K/I+ blue tholus containing a deeper-staining tube and pronounced ocular chamber, Bacidia-type, 8-spored. Ascospores 12-23 × 3-6 μm, narrowly ellipsoidal to cylindric-ellipsoidal with rounded ends, (1-)3-septate, hyaline, thin-walled, smooth, without an epispore, gelatinous sheath or appendages. Chemistry: lichen products not detected by TLC.