Thallus: basal squamules to 2 mm diam., very variable, ± rounded and sparingly indented; lower surface often yellow- to orange-brown, K+ purplish towards the base, ± dispersed or occasionally forming ± compact cushions. Podetia 1-2 cm tall, yellow-grey to yellow-green; scyphi wide, gradually tapering to the base with only a short stalk, regular or ± deformed, densely squamulose, rarely proliferating from the margins; surface smooth or uneven, ± corticate throughout, with corticate, easily detached granules, microsquamules and scale-like plates in the upper part and within the scyphi. Soredia usually absent. Anamorph: conidia pycnidia, frequent, red, on the margin of scyphi. Teleomorph: ascomata apothecia, red, frequent. Chemistry: thallus C–, K–, Pd–, UV– (usnic and porphyrilic acids, zeorin).